Remote northern Peru June – July 2016 - looking for Yellow-browed Toucanet and White-masked Antbird

Published by David Milton (david.milton AT csiro.au)

Participants: Sandra Harding, David Milton, Miguel Lezama Ninancuro

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Following on from the July – August 2015 trip to search for many of our 30 special birds to celebrate 30 years of birding, we decided to return to Peru to look more of our targets. We contacted Miguel Lezama Ninancuro from Tanager Tours (info1@tanagertours.com, www.tanagertours.com) for a suggested itinerary for northern Peru for 2016. Our plan was to travel to northern Peru in April/May as this would be good for fruit-eating birds such as the toucanets and toucans as it would be late wet season. As it turned out we delayed the trip until July and went to Brazil for a 2 week boat-based trip in the Amazon in June before heading on to Peru. This was probably a good move as Miguel had a considerably wet time on the way back from Plataforma in June, which we happily avoided.

Our itinerary aimed to maximise new birds and to see 14 possible targets species. This included three side trips: to a tributary of the Maranon River (Yellow-browed Toucanet) where we walked in with our 5 horse team; the Scarlet-banded Barbet four-wheel drive trip to Plataforma; and the small plane flight into the northern Amazon (San Lorenzo) to see the White -masked Antbird. At the end of the full 8 week trip to Brazil and Peru together with our 2015 trips to Peru (remote south and Cusco/Manu Rd area) and Japan we successfully achieved the 30 birds from our target families.

In the end, we did exceptionally well bird-wise, seeing 613 species between us, including 134 new for us and 9 of our targets. The total included 34 endemics and such rarities as Yellow-browed Toucanet, White-masked Antbird, Ochre-fronted, Pale-billed, Rusty-tinged and Chestnut Antpittas, Short-tailed Ant-thrush and Large-footed Tapaculo. The success of the trip was due to Miguel’s excellent guiding and exceptional logistical skills that meant we had no delays or hiccups, even in remote and often trying conditions.

17 to 20 June Sincicap to Huamachuco, Molino to San Marcos

From Manaus, Brazil we flew to Lima then to Trujillo (LATAM Airlines) arriving in the evening on 16 June. We had a chicken and chips dinner in a restaurant near our hotel. In the morning Miguel, Vidal and Ciaro were ready in an AWD station wagon that could fit all the camping equipment and luggage with careful packing. It was cold in Trujillo compared to the Amazonian lowlands in Brazil so that when we gained altitude we had all our clothing layers on and we knew we were back in Peru.

The drive to Sincicap was through dry scrub where we found Russet-bellied Spinetail, Black-necked Woodpecker and Collared Warbling-finch. We drove out of the Sincicap valley and up into the Andes and stayed the night in Huamachuco. On day 2 we got up at 5:30 am to get to Molino for the Purple-backed Sunbeam, which we had good views of the following day. We briefly considered camping but given the temperature of 10 degs C plus wind chill factor we drove on to stay in the closest small town called Arichapampa an hour on from Molino as this was the only accommodation available. We had the morning back in Molino seeing the hummingbirds and Barron’s Spinetail (Line-cheeked) in the small amount of damp forest left along a gulley. Miguel successfully called in the White-cheeked Cotinga and we also saw Spot-throated Hummingbird and Tyrian Metaltail. On the way back from Molino, Vidal tested out the car going up the steep bull-dust layered dirt tracks. That night we stayed in San Marcos and enjoyed quinoa pancakes made by Ciaro for breakfast the next day (Sunday). Vidal left us in the evening and went ahead via Celendin to Leymebamba to arrange for our horses as we weren’t entirely sure what the next leg of the trip involved.

21 to 22 June Celendin and Leymebamba

The next morning on Monday we were up above the clouds in the dry scrub that was pretty but freezing cold. Outside San Marcos, we saw the Great Spinetail and then Gray-bellied Comet in Chonta where we had lunch. We watched Grey-bellied Comets in the flowers while sitting in our comfortable chairs by the river we had vegetables, chicken and cassava topped with a bright purple olive sauce. The next day, it was a beautiful morning (Day 5) as we climbed over the ridge and looking down into the Maranon Valley, where we birded in several spots along the road. We were lucky to get several excellent views of the Yellow-faced Parrotlet and Grey-winged Inca-finch in a small area of natural vegetation including interesting spiny cactuses that was being encroached by farming. Crossing the Maranon valley and driving further up into mountains we spent time around the tree-line in stunted vegetation full of flowers and good for hummingbirds. We arrived in Leymebamba and got ready to go on the trek with the horses the next day at 8:30 am.

22 to 26 June Jardin, Orfedon to Leymebamba

It was about an hour’s drive up to the end of the road where we would meet our horse team who were waiting for us, having left Leymembamba at 2 am. Vidal had made the arrangements for Lennis and 5 horses which included two that we could ride as necessary. I was really impressed with the horses that could step up from muddy bogs and climb steep slippery rock tracks. They had me enthralled. Dave and I did ride till morning-tea but decided we preferred to walk the rest of the trail. It was a new birding experience for us and for everyone as the Yellow-browed Toucanet had not been searched for from this direction before by Miguel. When we reached Jardin, a small hut as the sun set, we were happy to have our tent put up inside the hut as we needed our thermal underwear to keep warm. We had great views of the Grey-throated Mountain-toucan just before the end of the day. Day 7 we walked further down the valley trying to locate the GPS fix that Miguel had of a previous toucanet sighting. The vigorous walking was made more challenging by the thigh-deep mud. Unlike the rest of the group, we had stuck to using our bushwalking boots so were trying to keep our balance on bits of vegetation and not waterlog the boots. We took the gumboot option on the way back.

The wet bromeliad forest around Jardin (2900m) was replaced with more wet tropical forest (wilderness photo status) with some cecropia trees which we thought should be suitable for attracting toucanets. Our next camp at Orfedon was an open grassy area on a shelf above the fast-flowing river and to get there we crossed a wooden bridge/walkway high up over a river. This was not something my horse at home could even contemplate crossing. We tried a muddy trail back on the other side of the river to see if we could find a place that was more open and where the river noise was not as loud. In the meantime, Ciaro saw a Yellow-browed Toucanet fly over at camp while we were away. That was the closest anyone got to the Yellow-browed Toucanet. We stayed another night at Orfedon to maximize our chances and did get really close excellent views of an Emerald Toucanet that we initially thought was a Yellow-browed. Despite the disappointment of not seeing the Yellow-browed Toucanet, we had some great bird watching seeing Coppery Metaltail, White-banded Tyrannulet, Inca Flycatcher, White-eared Solitaire, Chestnut-bellied Mountain-Tanager, White-collared Jay, Gray-breasted Wood-wren, Rufous-vented and Large-footed Tapaculos and Yellow-scarfed Tanager. Even though we did not see our target, we did get to some really remote and beautiful areas during the five day expedition.

27 June to 1 July Pomacochas, FAN, Moyobamba, Mishquiyacu to Bellavista

We left Leymebamba (Day 11) in light rain and drove back up the road to a small house that had some hummingbird feeders where we saw the Purple-throated Sunangel. We then returned to Leymembamba and continued north down the Leymebamba River valley. Miguel located the Maranon Crescentchest for us after lunch. In the evening, we visited the Huembo Ecological Centre and saw the Marvellous Spatuletail, a White-crowned Tapaculo came into the tape and a Little Woodstar was in flowers near the trail. We stayed at Pomachochas that night in order to be close to the Chido trail where we looked for Pale-billed Antpitta in the morning. The climb started in sunshine and lots of bird activity, but cloud quickly descended and the regular showers made birding hard – no luck with the Antpitta, not even a distant call. Travelling on to Fundo Alto Nieva (FAN) below Abra Particia, we birded a few spots along the road. We tried without luck for the Long-whiskered Owlet in the evening and again in the morning (4:30 am) and did get to hear it call. We saw the Speckle-chested Piculet in the morning (Day 13) in this habitat. We had a fantastic time at the feeders of Morro de Calzada in Aguas Verdes where we saw 17 different species of gorgeous hummingbirds, a special way to spend my birthday. We also saw Cinerous and Little Tinamous at a feeder. Ciaro made me a gluten-free birthday cake out of quinoa pancakes, really yummy for dinner at our lodge – Waqanki near Moyobamba. It was a good walk up to Mishquiyaquillo Mountains to see the Ash-throated Antwren the next day. Having walked through a coffee plantation on the way, we discovered to our dismay the next day that we had picked up a few chiggers. Driving out at night we saw Burrowing and Striped Owls between the lodge and Moyobamba town. The next day (day 15 - 1 July) we visited some small ponds and had good views of Comb Duck and stopped at the bridge to see the Oilbird, which we could just make out in the darkness given that our eyes were adjusted to bright sunlight. From there we drove down to Bellavista and got ready for our next adventure.

2 July to 4 July Plataforma to Bellavista

The 4-wheel drive track to Plataforma was in good condition but we were still amazed at the car’s (modified Toyota hilux ute) ability to climb up the steep slopes, and our driver and co-driver were obviously familiar with the road/track. We had a few stops to pick up White-necked Puffbird and Chestnut-eared and Ivory-billed Aracari. The track deteriorates into serious red mud in flatter areas where there can be a few alternative tracks to choose from. We arrived in the afternoon in good time to look for the Scarlet-banded Barbet and we had good views of 5 birds. It was a relief to see the target on the first afternoon. With generator on there was TV for several families and we could have an electric light, just had to take the bulb out to turn it off.

We were constantly looking at the clouds and speculating about rain while we were in Plataforma, but it kept away and walking was possible on the muddy tracks. We explored further along the ridge track the next morning and learnt about a horse that had fallen off the track from two small boys who found some more barbets for us. The forest is becoming more fragmented as agriculture (firstly coffee bushes) creeps onward. On a side track in good forest, a Short-tailed Antthrush crossed the track – nice to see. The drive back from Plataforma went well with just two small stops, one to fix an electrical problem with the car and the second to pay some road workers to remove trees that had been felled to block the track.

We walked around Bellavista in the afternoon that we arrived back, relaxing and trying to find somewhere to have coffee and in the end it was easier to have an ice cream. Miguel found an upmarket restaurant for dinner that had an outdoor dining area overlooking the town. We had the barbeque spare ribs, which was really tasty.

5 July to 9 July Yurimaguas to San Lorenzo to Yurimaguas

Day 19 began with a second quick look at the dry forest at quebrada Upaquigua, which turned up Rufous-capped Nunlet, then on into the La Escalera Mountains (tunnels) to see pretty Cobalt-winged and Rose-fronted Parakeets feeding on small berries near a burst water pipe. After much searching we found a feeding flock with Dotted Tanager. We made the most of our time at Koepeke’s Hermit Aconabekh where as well as hummingbird feeders (for Koepeke’s hermit) there was an army ant swarm in the vicinity. This produced White-plumed Antbird, Scale-backed Antbird, Chesnut-tailed Antbird and Black-faced Antthrush. The manager here was very helpful showing us a black and green striped frog and the roost of a Crested Owl. There seemed lots of potential in this forest particularly having active army ants.

It would be good to get back into the hot Amazon the following morning. After a spectacular small plane flight over the rainforest and swampy palm forests, we arrived in San Lorenzo (Day 20). As there were five of us we had a full plane and it left Yurimaguas as soon as the weather permitted. After landing in San Lorenzo, we quickly loaded up two toot-toots and went to Miguel’s known patch of forest for the White-masked Antbird. There are only one or two cars in San Lorenzo and everyone only uses toot-toots. This made a lot of sense with the extremely muddy roads. In order to leave town, you have to make a payment to a police guard to open the road. After that, we were into the forest along a small well-used logging trail. Very few people know about this site and there is no taping. Miguel had been there four times before and had learnt how to maximise our chances of seeing the birds in the dense undergrowth. Luckily, one Antbird responded to Miguel’s calls just on nightfall, giving us a brief view as it shot around in the dark forest.

Ciaro was kept busy making us our breakfasts and packed lunches each morning to eat in the forest, we were tired enough just getting up early. We were in the forest at dawn the next day (5:15), but it was raining hard and we sheltered in a small hut with chooks for a few hours in the middle of the day. A few hours of difficult walking in thick clinging mud along the road back to town provided a Black-banded Crake, so the ice cream back in town felt deserved. The next day (Day 22) we explored up a side river upstream from San Lorenzo in a small canoe with outboard to a new patch of forest. This required careful negotiation with land holders as it not good for one family to be seen as overly benefiting from outsiders compared to the rest of the village. As well as Canary-winged Parakeet, Dusky-billed Parrotlet, White-eared Jacamar, Western Striped and White-crowned Manakins and Cinereous Mourner, we had excellent views of two White-masked Antbirds. It was a stunning sighting and I felt really privileged.

We had a quick hour in the forest at the original site the next morning and saw Rufous-tailed Flatbill and had great views of the Black-necked Red Cotinga. Eight passengers and a slightly larger plane were ready at 12 PM for the flight back to Yurimaguas. so it was a late lunch by the road while driving back to Moyobamba. A brief stop was required in Tarapoto to pick up our table which had been accidently left behind in Plataforma

10 July to 13 July Moyobamba, FAN to Abra Patricia

We were keen to pick up a few antpittas and go back to look for the Long-whiskered Owlet and this was possible given we were ahead of our schedule. We stayed at Waqanki lodge and birded the lodge trail then headed back towards FAN. We stopped to look for the Point-tailed Palmcreeper along the road north of Rioja, a bird we had always wanted to see. At a lookout half way up to Abra Patricia, a flock of White-capped tanagers flew over - stunning birds. At Afluentes, the shop owner’s son had installed new hummingbird feeders after Miguel’s encouragement. These had brought in a lot of birds, but did have some harder to see species and we saw an Ecuadorian Piedtail.

It was good to be back at FAN (Day 24). Miguel decided the best chance for the Long-whiskered Owlet would be in the early evening, which proved to be the case. It came in beautifully and we were so lucky. We also tracked down the Cinnamon Screech Owl that we had also missed on the previous visit. Monday was a two antpitta day - Rusty-tinged and Ochre-fronted Antpittas. Also at FAN we enjoyed a Chestnut-breasted Wren and we rewarded the FAN guide with a tip. From FAN, we drove up the hill to Owlet Lodge (Abra Patricia) notable for the hummingbird feeders and other goodies. On the way, we had a brief view of a Chestnut Antpitta in secondary habitat near the road and spent the late afternoon looking for White-faced Nunbird. Some time looking for feeding flocks along the road yielded Red-billed Parrot, Equatorial Graytail and Green-and-black Fruiteater.

Over dinner that night we reviewed the remaining itinerary to see what we could pick back up as we drove west from Abra Patricia. We had time to go back and look for particular specialities so considering the options we wanted to look for the Pale-billed Antpitta on the Chido trail again. We got to the start of the Chido trail at 7 am (Day 27) and walked up to where we heard it across a valley in half an hour, as were now getting much fitter. I had a good view and Dave briefly saw it by eye as it walked away. This was disappointing for him, so back on the trail Miguel heard another one so we went into the bamboo and played the tape. Suddenly it was there, did not call but sat only 2 m away giving fantastic views. This was really worth changing the plans and it was only 9:30 am.

On the drive west to Jaen, we caught up Little Inca Finch in the dry forest along the middle Maranon River, but it was too hot at this time of day for much else.

14 July to 16 July Bosque Frejolillo, Olmos to Chiclayo

On Thursday, 14th we tried a new area – Bosque Yanahuanca - a small municipal forest one hour out of Jaen which was supposed to be a good site for Red-crested Finch and Maranon Spinetail. Arriving at dawn, we disturbed Scrub Nightjar and Lesser Nighthawk in the dry scrub in the dawn light. The park turned out to be very successful place to visit as we found both Maranon Spinetail and Red-crested Finch as well as many other species typical of that habitat. We even had fresh cow’s milk from the manager for breakfast and took lots of photos of a pet mongoose on a chain.

Arriving late at Bosque Frejolillo (White-winged Guan forest) (8 PM), we went looking for the Buff-fronted Owl that Miguel knew had been seen along the trail up the valley. We did not see or hear the owl, but did see the West Peruvian Screech-owl while Vidal and Ciaro set up our tent and cooked dinner. This was easy camping for us, but I’m sure Vidal and Ciaro must have been looking forward to having some down time after almost 30 days on the road. The walk up the dry river canyon the next day (Friday 15th) proved productive as we had several views of the Guan and Red-masked Parakeet. We also tracked down Short-tailed Woodstar, Tumbes Hummingbird, Guayaquil Woodpecker and Whooping Motmot. Our last day of the trip was at Bosque Pomac and we found all the targets reasonably quickly – Peruvian Plantcutter, Rufous Flycatcher, Cinereous Finch and Tumbes Swallow and caught up with Coastal Miner. We had a quick trip to the beach to see some gulls to add to our trip list and then it was time for the airport. After two, month-long trips together it felt like leaving family as we hugged and said our goodbyes. We had seen an amazing 610 species on the trip, been taken to remote, spectacular places and had several very enjoyable birding adventures, while being expertly-guided and looked after by Miguel, Vidal and Ciaro.

We departed from Chiclayo at 5:35 PM Saturday 16 to arrive in Lima to stay at the Manhattan Inn hotel near the airport where we relaxed for a day taking an international flight the following afternoon and arrived in Brisbane on Tuesday 19th.

Species Lists

Bird list (all seen)

Cinereous Tinamou Crypturellus cinereus
Little Tinamou Crypturellus soui
Torrent Duck Merganetta armata
Comb Duck Sarkidiornis sylvicola
Yellow-billed Teal Anas flavirostris
Masked Duck Nomonyx dominicus
Ruddy Duck Oxyura jamaicensis
Speckled Chachalaca Ortalis guttata
Sickle-winged Guan Charmaepetes goudotii
Andean Guan Penelope montagnii
White-winged Guan Penelope albipennis
Least Grebe Tachybaptus dominicus
Pied-billed Grebe Podilymbus podiceps
Andean Ibis Theristicus branickii
Puna Ibis Plegadis ridgwayi
Fasciated Tiger Heron Tigrisoma fasciatum
Black-crowned Night Heron Nycticorax nycticorax
Striated Heron Butorides striata
Western Cattle Egret Bubulcus ibis
Great Egret Ardea alba
Snowy Egret Egretta thula
Peruvian Pelican Pelecanus thagus
Neotropic Cormorant Phalacrocorax brasilianus
Turkey Vulture Cathartes aura
Black Vulture Coragyps atratus
Andean Condor Vultur gryphus
Swallow-tailed Kite Elanoides forficatus
Double-toothed Kite Harpagus bidentatus
Plumbeous Kite Ictinia plumbea
Snail Kite Rostrhamus sociabilis
Slate-colored Hawk Buteogallus schistaceus
Barred Hawk Morphnarchus princeps
Roadside Hawk Rupornis magnirostris
Harris's Hawk Parabuteo unicinctus
Variable Hawk Geranoaetus polyosoma
Black-chested Buzzard-Eagle Geranoaetus melanoleucus
White-throated Hawk Buteo albigula
Zone-tailed Hawk Buteo albonotatus
Russet-crowned Crake Laterallus viridis
Black-banded Crake Laterallus fasciatus
Grey-necked Wood Rail Aramides cajaneus
Blackish Rail Pardirallus nigricans
Plumbeous Rail Pardirallus sanguinolentus
Common Gallinule Gallinula galeata
Andean Coot Fulica ardesiaca
Peruvian Thick-knee Burhinus superciliaris
Black-necked Stilt Himantopus mexicanus
Andean Lapwing Vanellus resplendens
Pied Plover Hoploxypterus cayanus
Wattled Jacana Jacana jacana
Puna Snipe Gallinago andina
Andean Gull Chroicocephalus serranus
Grey-headed Gull Chroicocephalus cirrocephalus
Belcher's Gull Larus belcheri
Kelp Gull Larus dominicanus
Rock Dove Columba livia
Band-tailed Pigeon Patagioenas fasciata
Maranon Pigeon Patagioenas oenops
Plumbeous Pigeon Patagioenas plumbea
Plain-breasted Ground Dove Columbina minuta
Ruddy Ground Dove Columbina talpacoti
Ecuadorian Ground Dove Colombina buckleyi
Croaking Ground Dove Columbina cruziana
Blue Ground Dove Claravis pretiosa
White-tipped Dove Leptotila verreauxi
White-throated Quail-Dove Zentrygon frenata
Eared Dove Zenaida auriculata
West Peruvian Dove Zenaida meloda
Greater Ani Crotophaga major
Smooth-billed Ani Crotophaga ani
Groove-billed Ani Crotophaga sulcirostris
Squirrel Cuckoo Piaya cayana
West Peruvian Screech Owl Megascops roboratus
Cinnamon Screech Owl Megascops petersoni
Band-bellied Owl Pulsatrix melanota
Pacific Pygmy Owl Glaucidium peruanum
Long-whiskered Owlet Xenoglaux loweryi
Burrowing Owl Athene cunicularia
Striped Owl Pseudoscops clamator
Oilbird Steatornis caripensis
Lesser Nighthawk Chordeiles acutipennis
Anthony's Nightjar Nyctidromus anthonyi
Chestnut-collared Swift Streptoprocne rutila
White-collared Swift Streptoprocne zonaris
Pale-rumped Swift Chaetura egregia
Grey-rumped Swift Chaetura cinereiventris
Short-tailed Swift Chaetura brachyura
White-tipped Swift Aeronautes montivagus
Andean Swift Aeronautes andecolus
Neotropical Palm Swift Tachornis squamata
Pale-tailed Barbthroat Threnetes leucurus
Green Hermit Phaethornis guy
White-bearded Hermit Phaethornis hispidus
Great-billed Hermit Phaethornis malaris
Tawny-bellied Hermit Phaethornis syrmatophorus
Koepcke's Hermit Phaethornis koepckeae
Reddish Hermit Phaethornis ruber
Black-throated Hermit Phaethornis atrimentalis
Green-fronted Lancebill Doryfera ludovicae
Grey-breasted Sabrewing Campylopterus largipennis
White-necked Jacobin Florisuga mellivora
Brown Violetear Colibri delphinae
Green Violetear Colibri thalassinus
Sparkling Violetear Colibri coruscans
Black-throated Mango Anthracothorax nigricollis
Rufous-crested Coquette Lophornis delattrei
Wire-crested Thorntail Discosura popelairii
Blue-tailed Emerald Chlorostilbon mellisugus
Fork-tailed Woodnymph Thalurania furcata
Golden-tailed Sapphire Chrysuronia oenone
Tumbes Hummingbird Leucippus baeri
Spot-throated Hummingbird Leucippus taczanowskii
White-bellied Hummingbird Amazilia chionogaster
Amazilia Hummingbird Amazilia amazilia
Andean Emerald Amazilia franciae
Sapphire-spangled Emerald Amazilia lactea
Speckled Hummingbird Adelomyia melanogenys
Ecuadorian Piedtail Phlogophilus hemileucurus
Gould's Jewelfront Heliodoxa aurescens
Fawn-breasted Brilliant Heliodoxa rubinoides
Violet-fronted Brilliant Heliodoxa leadbeateri
Chestnut-breasted Coronet Boissonneaua matthewsii
Shining Sunbeam Aglaeactis cupripennis
Purple-backed Sunbeam Aglaeactis aliciae
Bronzy Inca Coeligena coeligena
Collared Inca Coeligena torquata
Violet-throated Starfrontlet Coeligena violifer
Rainbow Starfrontlet Coeligena iris
Sword-billed Hummingbird Ensifera ensifera
Great Sapphirewing Pterophanes cyanopterus
Giant Hummingbird Patagona gigas
Amethyst-throated Sunangel Heliangelus amethysticollis
Purple-throated Sunangel Heliangelus viola
Royal Sunangel Heliangelus regalis
Sapphire-vented Puffleg Eriocnemis luciani
Emerald-bellied Puffleg Eriocnemis aline
Greenish Puffleg Haplophaedia aureliae
Rufous-vented Whitetip Urosticte ruficrissa
Booted Racket-tail Ocreatus underwoodii
Black-tailed Trainbearer Lesbia victoriae
Green-tailed Trainbearer Lesbia nuna
Tyrian Metaltail Metallura tyrianthina
Coppery Metaltail Metallura theresiae
Black Metaltail Metallura phoebe
Grey-bellied Comet Taphrolesbia griseiventris
Long-tailed Sylph Aglaiocercus kingii
Marvelous Spatuletail Loddigesia mirabilis
Long-billed Starthroat Heliomaster longirostris
Oasis Hummingbird Rhodopis vesper
Peruvian Sheartail Thaumastura cora
Amethyst Woodstar Calliphlox amethystina
Purple-collared Woodstar Myrtis fanny
Short-tailed Woodstar Myrmia micrura
White-bellied Woodstar Chaetocercus mulsant
Little Woodstar Chaetocercus bombus
Golden-headed Quetzal Pharomachrus auriceps
Crested Quetzal Pharomachrus antisianus
Ecuadorian Trogon Trogon mesurus
Amazonian Trogon Trogon ramonianus
Blue-crowned Trogon Trogon curucui
Collared Trogon Trogon collaris
Masked Trogon Trogon personatus
American Pygmy Kingfisher Chloroceryle aenea
Green Kingfisher Chloroceryle americana
Ringed Kingfisher Megaceryle torquata
Whooping Motmot Momotus subrufescens
Broad-billed Motmot Electron platyrhynchum
White-eared Jacamar Galbalcyrhynchus leucotis
White-necked Puffbird Notharchus hyperrhynchus
Lanceolated Monklet Micromonacha lanceolata
Rufous-capped Nunlet Nonnula ruficapilla
Black-fronted Nunbird Monasa nigrifrons
White-fronted Nunbird Monasa morphoeus
Scarlet-crowned Barbet Capito aurovirens
Scarlet-banded Barbet Capito wallacei
Gilded Barbet Capito auratus
Versicolored Barbet Eubucco versicolor
Black-throated Toucanet Aulacorhynchus atrogularis
Yellow-browed Toucanet Aulacorhynchus huallagae
Chestnut-tipped Toucanet Aulacorhynchus derbianus
Lettered Aracari Pteroglossus inscriptus
Brown-mandibled Aracari Pteroglossus mariae
Chestnut-eared Aracari Pteroglossus castanotis
Golden-collared Toucanet Selenidera reinwardtii
Grey-breasted Mountain Toucan Andigena hypoglauca
Channel-billed Toucan Ramphastos vitellinus
Lafresnaye's Piculet Picumnus lafresnayi
Speckle-chested Piculet Picumnus steindachneri
Yellow-tufted Woodpecker Melanerpes cruentatus
Scarlet-backed Woodpecker Veniliornis callonotus
Bar-bellied Woodpecker Venilornis nigriceps
Little Woodpecker Veniliornis passerinus
Red-stained Woodpecker Veniliornis affinis
Golden-olive Woodpecker Colaptes rubiginosus
Crimson-mantled Woodpecker Colaptes rivolii
Black-necked Woodpecker Colaptes atricollis
Spot-breasted Woodpecker Colaptes punctigula
Andean Flicker Colaptes rupicola
Lineated Woodpecker Dryocopus lineatus
Crimson-bellied Woodpecker Campephilus haematogaster
Guayaquil Woodpecker Campephilus gayaquilensis
Mountain Caracara Phalcoboenus megalopterus
Northern Crested Caracara Caracara cheriway
Yellow-headed Caracara Milvago chimachima
American Kestrel Falco sparverius
White-winged Parakeet Brotogeris versicolurus
Cobalt-winged Parakeet Brotogeris cyanoptera
Red-billed Parrot Pionus sordidus
Blue-headed Parrot Pionus menstruus
Scaly-naped Parrot Amazona mercenaria
Southern Mealy Amazon Amazona farinosa
Dusky-billed Parrotlet Forpus modestus
Pacific Parrotlet Forpus coelestis
Yellow-faced Parrotlet Forpus xanthops
Rose-fronted Parakeet Pyrrhura roseifrons
Dusky-headed Parakeet Aratinga weddellii
Mitred Parakeet Psittacara mitratus
White-eyed Parakeet Psittacara leucophthalmus
Slender-billed Miner Geositta tenuirostris
Coastal Miner Geositta peruviana
Cream-winged Cinclodes Cinclodes albiventris
White-winged Cinclodes Cinclodes atacamensis
Pale-legged Hornero Furnarius leucopus
Pacific Hornero Furnarius cinnamomeus
Rusty-crowned Tit-Spinetail Leptasthenura pileata
White-chinned Thistletail Asthenes fuliginosa
Streak-throated Canastero Asthenes humilis
Many-striped Canastero Asthenes flammulata
Rufous Spinetail Synallaxis unirufa
Azara's Spinetail Synallaxis azarae
Dark-breasted Spinetail Synallaxis albigularis
Chestnut-throated Spinetail Synallaxis cherriei
Plain-crowned Spinetail Synallaxis gujanensis
Maranon Spinetail Synallaxis maranonica
Russet-bellied Spinetail Synallaxis zimmeri
Necklaced Spinetail Synallaxis stictothorax
Great Spinetail Siptornopsis hypochondriaca
Line-cheeked Spinetail Cranioleuca antisiensis
Baron's Spinetail Cranioleuca baroni
Russet-mantled Softtail Thripophaga berlepschi
Rufous-fronted Thornbird Phacellodomus rufifrons
Chestnut-backed Thornbird Phacellodomus dorsalis
Equatorial Greytail Xenerpestes singularis
Spotted Barbtail Premnoplex brunnescens
Pearled Treerunner Margarornis squamiger
Streaked Tuftedcheek Pseudocolaptes boissonneautii
Point-tailed Palmcreeper Berlepschia rikeri
Montane Foliage-gleaner Anabacerthia striaticollis
Rufous-necked Foliage-gleaner Syndactyla ruficollis
Buff-fronted Foliage-gleaner Philydor rufum
Olive-backed Foliage-gleaner Automolus infuscatus
Plain Xenops Xenops minutus
Streaked Xenops Xenops rutilans
Tyrannine Woodcreeper Dendrocincla tyrannina
White-chinned Woodcreeper Dendrocincla merula
Olivaceous Woodcreeper Sittasomus griseicapillus
Wedge-billed Woodcreeper Glyphorynchus spirurus
Black-banded Woodcreeper Dendrocolaptes picumnus
Straight-billed Woodcreeper Dendroplex picus
Buff-throated Woodcreeper Xiphorhynchus guttatus
Olive-backed Woodcreeper Xiphorhynchus triangularis
Streak-headed Woodcreeper Lepidocolaptes souleyetii
Montane Woodcreeper Lepidocolaptes lacrymiger
Fasciated Antshrike Cymbilaimus lineatus
Collared Antshrike Thamnophilus bernardi
Barred Antshrike Thamnophilus doliatus
Chapman's Antshrike Thamnophilus zarumae
Lined Antshrike Thamnophilus tenuepunctatus
Uniform Antshrike Thamnophilus unicolor
Mouse-colored Antshrike Thamnophilus murinus
Northern Slaty Antshrike Thamnophilus punctatus
Amazonian Antshrike Thamnophilus amazonicus
Rufous-capped Antshrike Thamnophilus ruficapillus
Dusky-throated Antshrike Thamnomanes ardesiacus
Bluish-slate Antshrike Thamnomanes schistogynus
Napo Stipple-throated Antwren Epinecrophylla haematonota
Foothill Antwren Epinecrophylla spodionota
Plain-throated Antwren Isleria hauxwelli
Stripe-chested Antwren Myrmotherula longicauda
White-flanked Antwren Myrmotherula axillaris
Grey Antwren Myrmotherula menetriesii
Ash-throated Antwren Herpsilochmus parkeri
Rusty-backed Antwren Formicivora rufa
Streak-headed Antbird Drymophila striaticeps
Rufous-rumped Antwren Terenura callinota
Blackish Antbird Cercomacra nigrescens
White-browed Antbird Myrmoborus leucophrys
Black-faced Antbird Myrmoborus myotherinus
Peruvian Warbling Antbird Hypocnemis peruviana
Silvered Antbird Sclateria naevia
Northern Chestnut-tailed Antbird Myrmeciza castanea
Southern Chestnut-tailed Antbird Myrmeciza hemimelaena
White-plumed Antbird Pithys albifrons
White-masked Antbird Pithys castaneus
Spot-backed Antbird Hylophylax naevius
Common Scale-backed Antbird Willisornis poecilinotus
Black-faced Antthrush Formicarius analis
Short-tailed Antthrush Chamaeza campanisona
Pale-billed Antpitta Grallaria carrikeri
Rusty-tinged Antpitta Grallaria przewalskii
Chestnut Antpitta Grallaria blakei
Ochre-fronted Antpitta Grallaricula ochraceifrons
Northern White-crowned Tapaculo Scytalopus atratus
Rufous-vented Tapaculo Scytalopus femoralis
Large-footed Tapaculo Scytalopus macropus
Maranon Crescentchest Melanopareia maranonica
Elegant Crescentchest Melanopareia elegans
Wing-barred Piprites Piprites chloris
Sooty-headed Tyrannulet Phyllomyias griseiceps
Black-capped Tyrannulet Phyllomyias nigrocapillus
Yellow-crowned Tyrannulet Tyrannulus elatus
Forest Elaenia Myiopagis gaimardii
Pacific Elaenia Myiopagis subplacens
Yellow-bellied Elaenia Elaenia flavogaster
Small-billed Elaenia Elaenia parvirostris
Sierran Elaenia Elaenia pallatangae
Southern Beardless Tyrannulet Camptostoma obsoletum
White-throated Tyrannulet Mecocerculus leucophrys
White-tailed Tyrannulet Mecocerculus poecilocercus
White-banded Tyrannulet Mecocerculus stictopterus
Black-crested Tit-Tyrant Anairetes nigrocristatus
Yellow-billed Tit-Tyrant Anairetes flavirostris
Tufted Tit-Tyrant Anairetes parulus
Unstreaked Tit-Tyrant Uromyias agraphia
Torrent Tyrannulet Serpophaga cinerea
Mouse-colored Tyrannulet Phaeomyias murina
Rufous-headed Pygmy Tyrant Pseudotriccus ruficeps
Tawny-crowned Pygmy Tyrant Euscarthmus meloryphus
Grey-and-white Tyrannulet Pseudelaenia leucospodia
Mishana Tyrannulet Zimmerius villarejoi
Golden-faced Tyrannulet Zimmerius chrysops
Variegated Bristle Tyrant Pogonotriccus poecilotis
Marble-faced Bristle Tyrant Pogonotriccus ophthalmicus
Mottle-cheeked Tyrannulet Phylloscartes ventralis
Streak-necked Flycatcher Mionectes striaticollis
Olive-striped Flycatcher Mionectes olivaceus
Ochre-bellied Flycatcher Mionectes oleagineus
Slaty-capped Flycatcher Leptopogon superciliaris
Inca Flycatcher Leptopogon taczanowskii
Flavescent Flycatcher Myiophobus flavicans
Olive-chested Flycatcher Myiophobus cryptoxanthus
Bran-colored Flycatcher Myiophobus fasciatus
White-eyed Tody-Tyrant Hemitriccus zosterops
Stripe-necked Tody-Tyrant Hemitriccus striaticollis
Pearly-vented Tody-Tyrant Hemitriccus margaritaceiventer
Black-throated Tody-Tyrant Hemitriccus granadensis
Cinnamon-breasted Tody-Tyrant Hemitriccus cinnamomeipectus
White-bellied Pygmy Tyrant Myiornis albiventris
Scale-crested Pygmy Tyrant Lophotriccus pileatus
Double-banded Pygmy Tyrant Lophotriccus vitiosus
Lulu's Tody-Flycatcher Poecilotriccus luluae
Rusty-fronted Tody-Flycatcher Poecilotriccus latirostris
Common Tody-Flycatcher Todirostrum cinereum
Yellow-browed Tody-Flycatcher Todirostrum chrysocrotaphum
Yellow-olive Flatbill Tolmomyias sulphurescens
Grey-crowned Flatbill Tolmomyias poliocephalus
Cinnamon Flycatcher Pyrrhomyias cinnamomeus
Cliff Flycatcher Hirundinea ferruginea
Black Phoebe Sayornis nigricans
Olive Tufted Flycatcher Mitrephanes olivaceus
Smoke-colored Pewee Contopus fumigatus
Tropical Pewee Contopus cinereus
Alder Flycatcher Empidonax alnorum
Vermilion Flycatcher Pyrocephalus rubinus
Rufous-tailed Tyrant Knipolegus poecilurus
Taczanowski's Ground Tyrant Muscisaxicola griseus
Rufous-naped Ground Tyrant Muscisaxicola rufivertex
Black-billed Shrike-Tyrant Agriornis montanus
White-tailed Shrike-Tyrant Agriornis albicauda
Streak-throated Bush Tyrant Myiotheretes striaticollis
Red-rumped Bush Tyrant Cnemarchus erythropygius
Rufous-webbed Bush Tyrant Polioxolmis rufipennis
Tumbes Tyrant Tumbezia salvini
Jelski's Chat-Tyrant Silvicultrix jelskii
Slaty-backed Chat-Tyrant Ochthoeca cinnamomeiventris
Rufous-breasted Chat-Tyrant Ochthoeca rufipectoralis
Brown-backed Chat-Tyrant Ochthoeca fumicolor
White-browed Chat-Tyrant Ochthoeca leucophrys
Piura Chat-Tyrant Ochthoeca piurae
Long-tailed Tyrant Colonia colonus
Short-tailed Field Tyrant Muscigralla brevicauda
Piratic Flycatcher Legatus leucophaius
Social Flycatcher Myiozetetes similis
Grey-capped Flycatcher Myiozetetes granadensis
Great Kiskadee Pitangus sulphuratus
Lesser Kiskadee Philohydor lictor
Lemon-browed Flycatcher Conopias cinchoneti
Baird's Flycatcher Myiodynastes bairdii
Streaked Flycatcher Myiodynastes maculatus
Boat-billed Flycatcher Megarynchus pitangua
Crowned Slaty Flycatcher Griseotyrannus aurantioatrocristus
Tropical Kingbird Tyrannus melancholicus
Rufous Flycatcher Myiarchus semirufus
Dusky-capped Flycatcher Myiarchus tuberculifer
Short-crested Flycatcher Myiarchus ferox
Brown-crested Flycatcher Myiarchus tyrannulus
Rufous-tailed Flatbill Ramphotrigon ruficauda
Red-crested Cotinga Ampelion rubrocristatus
White-cheeked Cotinga Zaratornis stresemanni
Peruvian Plantcutter Phytotoma raimondii
Green-and-black Fruiteater Pipreola riefferii
Fiery-throated Fruiteater Pipreola chlorolepidota
Andean Cock-of-the-rock Rupicola peruvianus
Black-necked Red Cotinga Phoenicircus nigricollis
Spangled Cotinga Cotinga cayana
Screaming Piha Lipaugus vociferans
Grey-tailed Piha Snowornis subalaris
Purple-throated Fruitcrow Querula purpurata
Dwarf Tyrant-Manakin Tyranneutes stolzmanni
Western Striped Manakin Machaeropterus striolatus
Fiery-capped Manakin Machaeropterus pyrocephalus
Blue-crowned Manakin Lepidothrix coronata
Band-tailed Manakin Pipra fasciicauda
White-crowned Manakin Dixiphia pipra
Golden-headed Manakin Dixiphia erythrocephala
Black-tailed Myiobius Myiobius atricaudus
Ruddy-tailed Flycatcher Terenotriccus erythrurus
Masked Tityra Tityra semifasciata
Cinereous Mourner Laniocera hypopyrra
White-browed Purpletuft Iodopleura isabellae
Green-backed Becard Pachyramphus viridis
Barred Becard Pachyramphus versicolor
White-winged Becard Pachyramphus polychopterus
Black-and-white Becard Pachyramphus albogriseus
Rufous-browed Peppershrike Cyclarhis gujanensis
Slaty-capped Shrike-Vireo Vireolanius leucotis
Brown-capped Vireo Vireo leucophrys
Red-eyed Vireo Vireo olivaceus
Lemon-chested Greenlet Hylophilus thoracicus
Ashy-headed Greenlet Hylophilus pectoralis
Tawny-crowned Greenlet Hylophilus ochraceiceps
White-collared Jay Cyanolyca viridicyanus
Violaceous Jay Cyanocorax violaceus
White-tailed Jay Cyanocorax mystacalis
Inca Jay Cyanocorax yncas
Tumbes Swallow Tachycineta stolzmanni
Grey-breasted Martin Progne chalybea
Blue-and-white Swallow Notiochelidon cyanoleuca
Brown-bellied Swallow Notiochelidon murina
White-banded Swallow Atticora fasciata
Southern Rough-winged Swallo Stelgidopteryx ruficollis
Black-capped Donacobius Donacobius atricapilla
Fasciated Wren Campylorhynchus fasciatus
Thrush-like Wren Campylorhynchus turdinus
Grey-mantled Wren Odontorchilus branickii
Sepia-brown Wren Cinnycerthia olivascens
Peruvian Wren Cinnycerthia peruana
Coraya Wren Pheugopedius coraya
Speckle-breasted Wren Pheugopedius sclateri
Buff-breasted Wren Cantorchilus leucotis
Superciliated Wren Cantorchilus superciliaris
House Wren Troglodytes aedon
Mountain Wren Troglodytes solstitialis
Grey-breasted Wood Wren Henicorhina leucophrys
Bar-winged Wood Wren Henicorhina leucoptera
Chestnut-breasted Wren Cyphorhinus thoracicus
Tropical Gnatcatcher Polioptila plumbea
Long-tailed Mockingbird Mimus longicaudatus
Andean Solitaire Myadestes ralloides
White-eared Solitaire Entomodestes leucotis
Great Thrush Turdus fuscater
Chiguanco Thrush Turdus chiguanco
Glossy-black Thrush Turdus serranus
Plumbeous-backed Thrush Turdus reevei
Maranon Thrush Turdus maranonicus
Black-billed Thrush Turdus ignobilis
White-capped Dipper Cinclus leucocephalus
House Sparrow Passer domesticus
Paramo Pipit Anthus bogotensis
Lesser Goldfinch Spinus psaltria
Hooded Siskin Spinus magellanicus
Purple-throated Euphonia Euphonia chlorotica
Thick-billed Euphonia Euphonia laniirostris
Bronze-green Euphonia Euphonia mesochrysa
White-vented Euphonia Euphonia minuta
Orange-bellied Euphonia Euphonia xanthogaster
Rufous-bellied Euphonia Euphonia rufiventris
Blue-naped Chlorophonia Chlorophonia cyanea
Tropical Parula Setophaga pitiayumi
Citrine Warbler Myiothlypis luteoviridis
Black-crested Warbler Myiothlypis nigrocristata
Grey-and-gold Warbler Myiothlypis fraseri
Russet-crowned Warbler Myiothlypis coronata
Three-banded Warbler Basileuterus trifasciatus
Three-striped Warbler Basileuterus tristriatus
Slate-throated Whitestart Myioborus miniatus
Spectacled Whitestart Myioborus melanocephalus
Peruvian Meadowlark Sturnella bellicosa
Russet-backed Oropendola Psarocolius angustifrons
Crested Oropendola Psarocolius decumanus
Solitary Cacique Cacicus solitarius
Yellow-rumped Cacique Cacicus cela
Yellow-tailed Oriole Icterus mesomelas
White-edged Oriole Icterus graceannae
Orange-backed Troupial Icterus croconotus
Epaulet Oriole Icterus cayanensis chrysocepha
Shiny Cowbird Molothrus bonariensis
Scrub Blackbird Dives warczewiczi
Oriole Blackbird Gymnomystax mexicanus
Bananaquit Coereba flaveola
Rufous-collared Sparrow Zonotrichia capensis
Yellow-browed Sparrow Ammodramus aurifrons
Tumbes Sparrow Rhynchospiza stolzmanni
Orange-billed Sparrow Arremon aurantiirostris
Black-capped Sparrow Arremon abeillei
Chestnut-capped Brush Finch Arremon brunneinucha
White-browed Brush Finch Arremon torquatus
Yellow-breasted Brush Finch Atlapetes latinuchus
Rufous-eared Brush Finch Atlapetes rufigenis
White-winged Brush Finch Atlapetes leucopterus
White-headed Brush Finch Atlapetes albiceps
Bay-crowned Brush Finch Atlapetes seebohmi
Common Bush Tanager Chlorospingus flavopectus
Yellow-whiskered Bush Tanager Chlorospingus parvirostris
Yellow-throated Bush Tanager Chlorospingus flavigularis
Ashy-throated Bush Tanager Chlorospingus canigularis
Black-faced Tanager Schistochlamys melanopis
Magpie Tanager Cissopis leverianus
White-capped Tanager Sericossypha albocristata
Rufous-crested Tanager Creurgops verticalis
Black-capped Hemispingus Hemispingus atropileus
Superciliaried Hemispingus Hemispingus superciliaris
Oleaginous Hemispingus Hemispingus frontalis
Black-eared Hemispingus Hemispingus melanotis
Drab Hemispingus Hemispingus xanthophthalmus
Grey-hooded Bush Tanager Cnemoscopus rubrirostris
Rufous-chested Tanager Thlypopsis ornata
Buff-bellied Tanager Thlypopsis inornata
Yellow-crested Tanager Tachyphonus rufiventer
Fulvous-crested Tanager Tachyphonus surinamus
White-lined Tanager Tachyphonus rufus
Masked Crimson Tanager Ramphocelus nigrogularis
Huallaga Tanager Ramphocelus melanogaster
Silver-beaked Tanager Ramphocelus carbo
Blue-grey Tanager Thraupis episcopus
Palm Tanager Thraupis palmarum
Blue-capped Tanager Thraupis cyanocephala
Blue-and-yellow Tanager Thraupis bonariensis
Hooded Mountain Tanager Buthraupis montana
Lacrimose Mountain Tanager Anisognathus lacrymosus
Scarlet-bellied Mountain Tanage Anisognathus igniventris
Blue-winged Mountain Tanager Anisognathus somptuosus
Grass-green Tanager Chlorornis riefferii
Buff-breasted Mountain Tanager Dubusia taeniata
Chestnut-bellied Mountain Tanager Delothraupis castaneoventris
Yellow-throated Tanager Iridosornis analis
Yellow-scarfed Tanager Iridosornis reinhardti
Orange-eared Tanager Chlorochrysa calliparaea
Turquoise Tanager Tangara mexicana
Paradise Tanager Tangara chilensis
Green-and-gold Tanager Tangara schrankii
Golden Tanager Tangara arthus
Saffron-crowned Tanager Tangara xanthocephala
Golden-eared Tanager Tangara chrysotis
Flame-faced Tanager Tangara parzudakii
Yellow-bellied Tanager Tangara xanthogastra
Dotted Tanager Tangara varia
Bay-headed Tanager Tangara gyrola
Golden-naped Tanager Tangara ruficervix
Metallic-green Tanager Tangara labradorides
Blue-browed Tanager Tangara cyanotis
Blue-necked Tanager Tangara cyanicollis
Masked Tanager Tangara nigrocincta
Beryl-spangled Tanager Tangara nigroviridis
Blue-and-black Tanager Tangara vassorii
Silver-backed Tanager Tangara viridicollis
Swallow Tanager Tersina viridis
Black-faced Dacnis Dacnis lineata
Blue Dacnis Dacnis cayana
Short-billed Honeycreeper Cyanerpes nitidus
Purple Honeycreeper Cyanerpes caeruleus
Green Honeycreeper Chlorophanes spiza
Guira Tanager Hemithraupis guira
Yellow-backed Tanager Hemithraupis flavicollis
Cinereous Conebill Conirostrum cinereum
Blue-backed Conebill Conirostrum sitticolor
Capped Conebill Conirostrum albifrons
Moustached Flowerpiercer Diglossa mystacalis
Black-throated Flowerpiercer Diglossa brunneiventris
White-sided Flowerpiercer Diglossa albilatera
Bluish Flowerpiercer Diglossa caerulescens
Masked Flowerpiercer Diglossa cyanea
Red Pileated Finch Coryphospingus cucullatus
Peruvian Sierra Finch Phrygilus punensis
Plumbeous Sierra Finch Phrygilus unicolor
Ash-breasted Sierra Finch Phrygilus plebejus
Cinereous Finch Piezorina cinerea
Grey-winged Inca Finch Incaspiza ortizi
Buff-bridled Inca Finch Incaspiza laeta
Little Inca Finch Incaspiza watkinsi
Collared Warbling Finch Poospiza hispaniolensis
Bright-rumped Yellow Finch Sicalis uropigyalis
Saffron Finch Sicalis flaveola
Wedge-tailed Grass Finch Emberizoides herbicola
Blue-black Grassquit Volatinia jacarina
Yellow-bellied Seedeater Sporophila nigricollis
Drab Seedeater Sporophila simplex
Chestnut-bellied Seedeater Sporophila castaneiventris
Chestnut-bellied Seed Finch Oryzoborus angolensis
Black-billed Seed Finch Oryzoborus atrirostris
Band-tailed Seedeater Catamenia analis
Plain-colored Seedeater Catamenia inornata
Dull-colored Grassquit Tiaris obscurus
Hepatic Tanager Piranga hepatica
Summer Tanager Piranga rubra
Scarlet Tanager Piranga olivacea
White-winged Tanager Piranga leucoptera
Golden Grosbeak Pheucticus chrysogaster
Slate-colored Grosbeak Saltator grossus
Buff-throated Saltator Saltator maximus
Greyish Saltator Saltator coerulescens
Golden-billed Saltator Saltator aurantirostris
Streaked Saltator Saltator striatipectus

Mammal list

Dusky titi monkey
Red squirrel
Tyra
Grey and white mongoose
Grey and red squirrel
Grey mongoose