Panama - September 29th - October 25th 2015

Published by Greg Roberts (friarbird.roberts AT

Participants: Greg Roberts (leader, Australia), Euclides Campos (guide, Panama), Ketil Knudsen (Norway), Niels Poul Dreyer (Denmark), Taus Rasmussen (Denmark), Jeff Skevington (Canada), Jonathan Newman (United Kingdom), Bill Watson (Australia), Barbara De Witt (United



Greg Roberts -

Most of this extensive birding trip to Panama was organised by Birding Panama ( All the participants had previously birded Costa Rica and Colombia so we were focused on regional specialties and species that are difficult to find elsewhere. We had an extraordinarily successful trip with almost all key target birds seen. Almost 600 species were recorded with good birds including Black-eared Wood-Quail, Agami Heron, Crested Eagle, Plumbeous Hawk, Slaty-backed Forest-Falcon, Rufous-vented Ground-Cuckoo, Costa Rica Pygmy-Owl, Dusky Nightjar, Tooth-billed Hummingbird, Veraguan Mango, White-bellied Mountain-Gem, Snowcap, Black-bellied Hummingbird, Pirre Hummingbird, Yellow-eared Toucanet, Blue-fronted Parrotlet, Lattice-tailed Trogon, Resplendent Quetzal, Tody Motmot, Barred Puffbird, White-whiskered Puffbird, Grey-cheeked Nunlet, Stripe-cheeked Woodpecker, Crimson-bellied Woodpecker, Sulphur-winged Parakeet, Spot-crowned Barbet, Tody Motmot, Speckle-faced Antbird, Ocellated Antbird, Wing-banded Antbird, Black Antshrike, Speckled Antshrike, Black-crowned Antpitta, Streak-breasted Antpitta, Ochre-breasted Antpitta, Black-headed Ant-thrush, Sapayoa, Brown-billed Scythebill, Silvery-fronted Tapaculo, Streak-breasted Treehunter, Slaty-winged Foliage-gleaner, Beautiful Treerunner, Speckled Mourner, Double-banded Greytail, Yellow-green Tyrannulet, Choco Sirystes, Brownish Twistwing, Ochraceous Peewee, Dark Peewee, Russet-winged Schiffornis, Northern Schiffornis, Blue Cotinga, Sharpbill, Silvery-throated Jay, Sooty-headed Wren, Stripe-throated Wren, Slate-throated Gnatcatcher,Varied Solitaire, Pirre Warbler, Zeledonia (Wrenthrush), Connecticut Warbler, Green-naped Tanager, Blue-and-gold Tanager, Pirre Bush-Tanager,Viridian Dacnis, Orange-collared Manakin, Green Manakin, Yellow-green Finch and Black Oropendola.

I organised the itinerary in conjunction with Jose Carlos Garcia ( I can highly recommend the services of Birding Panama; this was a difficult trip with plenty of challenges but with the exception of a couple of inevitable hiccups, it was very well organised. We were guided by Euclides (Kilo) Campos, who was by any standard a highly capable and skilled guide, able to track down the most difficult skulkers and always willing to help. In the Darien we were also guided by Isaac Pizaro from the local Guna Indian community, who has extensive knowledge of the local avifauna; Isaac organised the on-the-ground logistics for our visit to Cerro Pirre. Others have commented in reports that Isaac's behaviour can be erratic; there is some truth to this but at the end of the day he came up with the goods.
A series of seven illustrated posts from the trip have been published on my blog (search “Panama” on

We visited major sites in the Darien in the east of the country near the Colombia border; in central Panama; and in the Chiriqui highlands in the west near the Costa Rica border. Trails were muddy and steep in places. Trips of this nature are not easy, especially with a large group. However we were fortunate with the weather, losing very little birding time to rain, and frequently overcast conditions kept temperatures in check to some extent. We had no mechanical difficulties although the bus could have been more comfortable.

The biggest challenge of the trip was always going to be the assault on Cerro Pirre in the Darien – a key and difficult-to-access site for many regional endemics and specialties. We had three full days in this area in addition to two travel days so hard decisions had to be made about how the time should best be utilised. Our base for this part of the trip was Rancho Frio, the headquarters of Darien National Park. We had three nights at Rancho Frio and two nights camping on Cerro Pirre, with our gear being transported up steep tracks by Isaac's excellent team of porters and other workers.

We spent the first of the three days in the lowland forests around Rancho Frio. This was a good move because we cleaned up most of the lowland and lower foothill specialties, allowing more time to look for mid-elevation species on our way up Cerro Pirre on the second day. The first camping night was at mid-elevation (640m) at camp site called Rancho Plastico. The group split for the second night, with some again camping at Plastico and others ascending to a ridge camp at 1100m. Some specialties are found only on the higher slopes but not everybody could camp up there, in part because of the physical challenges involved in the very steep climb and also because provisions for a second camping party were limited.

With the benefit of hindsight, I believe it would have been better for the whole group to have camped both nights at Plastico, with those wishing to go up to the ridge leaving early in the morning and returning in the afternoon. It would have made operations much easier logistically and prevented tensions that can arise when guides and other resources are split. While formidable and challenging, the climb to the top was not as dire as some reports suggest and can be done quite easily in a full day if participants are reasonably fit.


September 29. I arrived in Panama City following a 27-hour sojourn from Brisbane via Los Angeles. Overnight in the very nice and birdy Radisson Summit Hotel.

September 30. Birding in the forest which surrounds the hotel.

October 1. The first day of the 21-day tour but the day before our travelling began. Some of us who arrived early hired Kilo for a morning excursion to Chagres National Park, a nice area of lowland rainforest not far from the hotel. Good sightings included exceptionally close-up views of the endemic Yellow-green Tyrannulet. Blue Cotinga, Connecticut Warbler and White-whiskered Puffbird were seen.

October 2. Today we visited the famed Pipeline Road in Soberania National Park, also not far from the hotel. We spent the day walking the shaded road, scoring nicely with Spotted Antbird, Russet-winged Shiffornis and Streak-breasted Antpitta. An Agami Heron close to the track was an unexpected bonus.

October 3. We left the hotel early, crossing the Panama Canal and heading east along the Pan-American Road to the Bayano lowlands, where we had Black Antshrike in scrub by the Rio Mono. We birded some more scrub near Torti along the Rio Torti, finding Pacific Antwren and Double-banded Greytail. Lunch was at the Hotel Torti where we were entertained by an abundance of hummers at the feeders including Scaly-breasted Hummingbird, Snowy-bellied Hummingbird and Long-billed Starthroat. We packed small bags for our foray into the Darien and left our main luggage cases, laptops and the like in storage at the hotel to pick up on the way back; this was necessary because the travel arrangements for the Darien are not adequate to handle large bags. We headed further to east to Meteti, overnighting in the basic but pleasant Meteti Hotel.

October 4. We checked out some secondary scrub and grassland east of Metiti early in the morning, then had some very good birding at the eastern end of the Pan-American Road, especially roadside in forest patches within 15km of Yaviza, the busy port town where the road terminates. Here we saw Grey-cheeked Nunlet, Barred Puffbird, Spot-crowned Barbet and Black Oropendola, along with Geoffrey's Tamarin. We met Isaac Pizaro and loaded ourselves into a large motorised canoe for a 1.5-hour journey up the Rio Chucumaque to the town of El Real. After lunch we crowded into an ancient truck for a 10km drive to the edge of the forest. We then hiked 6km through the forest to Rancho Frio, the national park headquarters.

Our trek was interrupted by the sight of an imposing and much-wanted dark phase Crested Eagle – one of the most difficult neotropical raptors to see - perched high in a tree above the track. This was near the spot where a pair of Harpy Eagles had for several years nested successfully until earlier this year, when one of the adults was shot by locals; the people evidently were angry that their demands for money from visiting birders in a national park buffer zone were refused. After a long and eventful day, we arrived in the late afternoon at Rancho Frio, pleasantly located in a clearing in the forest beside a fast-running stream, ideal for bathing. Our baggage arrived on the backs of three horses. Accommodation was in a shared dormitory.

October 5. A day on the lowland and lower foothill forest trails around Rancho Frio. It started well with Viridian Dacnis and Choco Sirystes near the buildings. Soon after we had a pair of Sapayoa – the only one of the world's 234 bird families that I had not seen. In the same area we saw Slaty-winged Foliage-gleaner and Lemon-spectacled Tanager. A Speckled Mourner foraging mid-canopy was another species that is difficult to find elsewhere.

Late in the day, when we thought it couldn't get much better, Kilo was alarmed when he heard, close to the track, the sound of what he thought was the gnashing of tusks of potentially dangerous White-lipped Peccaries. Instead, out of the thick ground cover popped a Rufous-vented Ground-Cuckoo, which perched briefly on a log before fluttering into the undergrowth; unfortunately half the group missed the bird.

October 6. The porters had gone up the steep slopes of Cerro Pirre yesterday to set up our mid-elevation camp at Plastico. Early on our hike up the mountain we scored with Plumbeous Hawk - another difficult neotropical raptor. Then came a suite of highly desirable mid-elevation birds: Tody Motmot, Yellow-eared Toucanet, Dull-mantled Antbird, Black-and-yellow Tanager, White-ruffed Manakin and best of all, Wing-banded Antbird. This is probably the best place to see Wing-banded Antbird and our excellent views of 2 birds on the slopes below the trail were among the trip highlights.

The camp at Plastico was basic with small tents under the cover of a large plastic sheet. Unfortunately, sleeping mats we thought would be provided did not materialise, evidently due to a misunderstanding. Most of us were content enough to put up with the discomforts in the knowledge that getting a large group up this mountain was never going to be a fairy tale logistically. Kilo and the guys went the extra mile to meet various requests, to the point of compromising their own comfort. In the late afternoon during a steep walk down to a stream, we saw more Sapayoas.

October 7. This day presented new challenges because the group divided, with most heading uphill to another camp on the ridge at 1100m. Not everybody could or was able to go up to the ridge for various reasons (see discussion above) so some remained at Plastico for the second camping night. Those who ascended further were rewarded with the endemic Stripe-cheeked Woodpecker not far from camp, and further up with a feeding flock of Blue-fronted Parrotlets - a species normally seen only as a quick fly-by, if at all. Brown-billed Scythebill was another welcome addition to the list.

Higher up still, Cerro Pirre's specialties emerged in the form of Varied Solitaire and Pirre Hemispingus, while Tooth-billed Hummingbird and Violet-throated (Emerald) Toucanet put in appearances. At the summit, Pirre Hummingbird and Sooty-headed Wren showed nicely, while most of the group saw Pirre Warbler and Choco Tapaculo. Late in the day, some of us had crippling views of a nicely co-operative Black-crowned Antpitta – another species high on our wishlists - close to Plastico camp.

October 8. The good fortune of those at the top continued in the form of Black-eared Woodquail, Russet-crowned Quail-Dove and Beautiful Treerunner. Those who remained at Plastico made do with great views of Central American Pygmy-Owl, Ornate Hawk-Eagle and another Stripe-cheeked Woodpecker. All us eventually made our way slowly back down the steep trail to Rancho Frio in the afternoon. Not long before sunset, a fledgling Crested Eagle close to camp made quite a din as it begged for food from parents unseen. Around the camp, Crested Owl and Choco Screech-Owl were heard but not seen.

October 9. Our journey to Rancho Frio in reverse: hiking back through the forest, a truck ride to El Real, then the boat back to Yaviza, this time at low tide with good numbers of waterbirds feeding along the shores. We headed west to fetch our luggage at the Hotel Torti then moved on to our next destination: Burbayar Lodge in the Caribbean foothills of central-east Panama. This lodge had long been a primary destination for birders in Panama but in recent years, many have reported problems with booking accommodation.

October 10. We walked some of steep and muddy trails a few kilometres from the lodge in Nusagandi – a large forest reserve owned by local Indian communities. We saw Crimson-bellied Woodpecker, Tawny-capped Euphonia and Striped (Western) Woodhaunter along with our only large ant swarm of the trip, with attendants including several obliging Ocellated Antbirds.

October 11. Early in the morning we scoped a Plumbeous Hawk from the lodge on a slope across a valley. We visited a forested gully in another area near the lodge, seeing yet more Sapayoas (a total of 12-14 were seen during the trip) and Sulphur-rumped Tanager. Some of the group in the afternoon visited another site to connect with Speckled Antshrike.

October 12. A day largely in transit, heading west back across the Panama Canal to the resort town of El Valle, located in a volcanic crater in Panama's central highlands, where we stayed in the Anton Valley Hotel.

October 13. We visited some nice forest patches around Cerro Gaital and second-growth scrub near El Valle, seeing Pale-vented Thrush, Garden Emerald and Northern Schiffornis. With the benefit of hindsight we did not need to visit this site, but it was intended as back-up for key targets such as Black-crowned Antpitta and Rufous-vented Ground-Cuckoo, which we unexpectedly cleaned up in the Darien.

October 14. We continued west on a long drive to Las Lajas, a coastal site in north-west Panama, where after some effort along the main road to the beach we spotted the local specialty – a male Veraguan mango, another Panama endemic. We moved on through the city of David to the Chiriqui Highlands town of Volcan, where we booked into the delightful Hotel Dos Rio.

October 15. This morning early we headed to the Volcan foothills site of Cuesta de Piedra, a 30-minute drive from the hotel, where we ticked off regional specialties including Cherrie's Tanager and Costa Rica Brushfinch. Eye-ringed Flatbill, seen by some on Cerro Pirre, was present. We moved on to Volcan Lakes, seeing Orange-collared Manakin in remnant scrub and the distinctive Chiriqui race of Masked Yellowthroat.

October 16. We had the whole day on the upper slopes (between 1800 and 2500m) of Volcan Buru National Park along the Los Quetzales Trail. Here in the beautiful cloud forest we had a feast of specialties shared with neighbouring Costa Rica, where many are more difficult to find than in Panama. Early in the morning we had mixed flocks lower down including Scintillant Hummingbird, Black-cheeked Warbler and White-throated Mountain-Gem. Long-tailed Silky-Flycatcher was in the mix along with Volcano Hummingbird and Resplendent Quetzal higher up.

At the highest point of our walk we connected nicely with a Zeledonia, or Wrenthrush – likely to eventually have a family of its own – in the undergrowth; we were to snatch glimpses of a couple more as the day progressed. Silvery-fronted Tapaculo was another skulker that showed briefly at first, then very well on the track. Then came two much-wanted specialties in quick succession - a flock of 8-10 Silvery-throated Jays and an Ochraceous Peewee; both species are challenging to find elsewhere. The day was capped off by a vocal Costa Rica Pygmy-Owl tracked down after some effort.

October 17. We checked out of the hotel and moved to another scenic highlands tourist town – Boquete, where we booked into Boquete Tree-trek at 1400m. Along the road in the afternoon we saw several Dark Peewees and a flock of Sulphur-winged Parakeets, while a probable Maroon-chested Ground-Dove flew across the road, not to be found again.

October 18. We had a long (2.5 hours) drive to Fortuna Reserve on the Continental Divide that separates the Pacific and Caribbean slopes of western Panama. We began on the Pacific side at about 1000m with good numbers of hummingbirds including White-bellied Mountain-Gem, Snowcap and Black-bellied Hummingbird; the latter two were unexpected but most welcome. Another vocal Costa Rican Pygmy-Owl attracted large numbers of hummers and other birds. Blue-and-gold Tanager, a Panama endemic, showed nicely and was not uncommon along the road. On the Caribbean side, we headed down a forest trail and scored stunning views of two highly desirable birds: Lattice-tailed Trogon and Ochre-breasted Antpitta. On another trail from where the Caribbean Sea glistened in the distance, a Black-headed Ant-thrush showed nicely.

October 19. In the morning, we birded the road close to the hotel, adding Philadelphia Vireo to the list, before moving on to David and east to the town of San Felix, where we booked into a basic hotel.

October 20. We were up early for a 1.5-hour drive northwards to the cloud forest of Cerro Santiago. It was dark when we arrived to the loud calling of a Dusky Nightjar by the road. We saw 1 or 2 birds fly by closely and Ketil managed to photograph one with the improbable aid of a smartphone torch. As dawn broke, we had several Streak-breasted Treehunters about along with our main target – Yellow-green Finch, another Panama endemic. Our birding success was followed by a long drive back to Panama City, where we again booked into the Radisson Summit.

October 21. A day of rest and contemplation. With the benefit of hindsight, I would not have organised things much differently. If we had known we were going to do so well in the Darien, we would not have gone to El Valle; we could have done with an extra night on Cerro Pirre, but the logistical challenges were substantial. Nor do I think we should have split the overnight camping into two groups for the second night on Cerro Pirre.

Unfortunately it happens sometimes with large group excursions that an individual may be more concerned with his or her personal comforts than with the interests of the group; that adds to the challenges of running an already difficult operation. However, we had outstanding success birdwise, making up for the relatively minor pitfalls and annoyances. Most of these were outside the control of what for the most part was an excellent and highly motivated group of experienced and enthusiastic birders. And many thanks to Kilo and Jose Carlos again for their wonderful work.

October 22. A dude visit to Panama Canal. Birding over, for the time being.

October 23. Another dude visit – to Casco Viejo, the Old City of Panama.

October 24. Depart Panama City.

Species Lists

Common names and taxonomy follow The Clements Checklist of Birds of the World including updates to 2015, and The Birds of Panama – a Field Guide (Angehr & Dean 2010). No effort is made here to provide annotations for common and widespread species. [] = heard only. {} = not seen by me. References to Burbayar include various forest patches in the region in addition to the lodge. References to Volcan, Boquete and El Valle include various sites in the vicinity of those towns. Volcan Buru is the national park of that name. Reference to Pan American Rd includes sites along the road between Panama City and Metiti. The term “widespread” indicates the species was recorded at more than 4 sites. Full species lists for individual sites have been published by Jeff Skevington on e-bird; as the group was often separated, some entries on those lists do not necessarily tally with the list below.

Great Tinamou (2 Pipeline Rd, 1 Cerro Pirre; heard elsewhere eastern lowlands),
[Little Tinamou – Radisson Summit Hotel, Burbayar],
{Choco Tinamou – (1 mid-elevation Cerro Pirre)},
Black-bellied Whistling-Duck, Blue-winged Teal,
Grey-headed Chachalaca (widespread, common), Crested Guan (1 lower slopes Cerro Pirre),
Black Guan (1 Volcan Baru), [Marbled Wood-Quail – Rancho Frio],
{Black-eared Wood-Quail (1 Cerro Pirre ridge; others heard)},
[Black-breasted Wood-Quail – Cerro Santiago], [Spotted Wood-Quail – Volcan Baru, Volcan],
Least Grebe, Wood Stork, Magnificent Frigatebird, Brown Booby, Neotropic Cormorant,
Anhinga, Brown Pelican, Great Blue Heron, Cocoi Heron, Rufescent Tiger-Heron,
Agami Heron (1 Pipeline Rd), Great Egret, Snowy Egret, Little Blue Heron, Tricoloured Heron,
Cattle Egret, Green Heron (widespread), Striated Heron (1 Rancho Frio),
Capped Heron (a few from boat to El Real), Black-crowned Night-Heron,
Yellow-crowned Night-Heron (several from boat to El Real), American White Ibis, Green Ibis,
Black Vulture, Turkey Vulture, Lesser Yellow-headed Vulture (1 Las Lajas),
King Vulture (a few Pan-American Rd, Burbayar, south of David),
Osprey, [Grey-headed Kite – Chagres], White-tailed Kite (1 near Santiago),
Hook-billed Kite (2 from boat to El Real), Pearl Kite (a few singles roadside),
Double-toothed Kite (1 Torti), Mississippi Kite (small flocks migrating eastern lowlands),
Plumbeous Kite (a few Pipeline Rd), Black-collared Hawk (1 from boat to El Real),
Crane Hawk (1 from boat to El Real, 1 Burbayar),
Common Black-Hawk (sparse, widespread), [Great Black-Hawk – Cerro Pirre],
Plumbeous Hawk (1 Rancho Frio, 1 Burbayar),
Savanna Hawk (1 east of Metiti), White Hawk (several eastern lowlands),
Semiplumbeous Hawk (1 Burbayar), Barred Hawk (1 Boquete), Roadside Hawk,
Broad-winged Hawk (flocks totalling tens of thousands migrating, especially from boat to El Real),
Grey-lined Hawk (widespread in small numbers), Short-tailed Hawk (1 Rio Mono),
Swainson's Hawk (a few western highlands), Zone-tailed Hawk (1 Torti),
Red-tailed Hawk ( a couple western highlands),
Crested Eagle (2 Rancho Frio),
Black Hawk-Eagle (1 Cerro Pirre),
Ornate Hawk-Eagle (1 Cerro Pirre; heard Pipeline Rd; singles Boquete, Fortuna),
Black-and-white Hawk-Eagle (a couple Cerro Pirre),
[White-throated Crake – scattered sites], [Grey-breasted Crake – east of Metiti],
Grey-necked Wood-Rail (several from boat to El Real; 1 Volcan; heard Pipeline Rd),
Purple Gallinule, Common Moorhen, Southern Lapwing, Grey Plover,
American Oystercatcher, Northern Jacana, Wattled Jacana,
Spotted Sandpiper, Solitary Sandpiper, Willet, Lesser Yellowlegs, Whimbrel,
Semipalmated Sandpiper, Least Sandpiper, Pectoral Sandpiper,
Black Tern, Common Tern, Royal Tern,
Rock Dove, Pale-vented Pigeon (widespread), Scaled Pigeon (a few Pipeline Rd, Burbayar),
Band-tailed Pigeon (small numbers western highlands),
Short-billed Pigeon (a few Chagres, Rancho Frio, Burbayar),_
{Russet-crowned Quail-Dove (several Cerro Pirre ridge)},
{Violaceous Quail-Dove (a couple mid-elevation Cerro Pirre)}, [Ruddy Quail-Dove – Rancho Frio],
Plain-breasted Ground-Dove (a few Boquete), Ruddy Ground-Dove (common),
Blue Ground-Dove (sparse, widespread), White-tipped Dove (common eastern lowlands),
Grey-headed Dove (a few Cerro Pirre foothills, Burbayar), Grey-chested Dove (1 Chagres),
Squirrel Cuckoo (widespread), Yellow-billed Cuckoo (1 Chagres),
Black-billed Cuckoo (1 Chagres), Striped Cuckoo (2 east of Metiti),
Rufous-vented Ground-Cuckoo (1 at Rancho Frio was a trip highlight),
Greater Ani (common eastern lowlands), Smooth-billed Ani, Groove-billed Ani,
[Tropical Screech-Owl – Volcan],
[Verminculated (Choco) Screech-Owl – Rancho Frio, Plastico camp on Cerro Pirre],
[Crested Owl – Rancho Frio], [Spectacled Owl – Rancho Frio],
Costa Rican Pygmy-Owl (1 Volcan Buru; 1 Fortuna; heard Boquete),
Central American Pygmy-Owl (1 Cerro Pirre Plastico, others calling; 1 Burbayar),
[Mottled Owl – Rancho Frio], Striped Owl (1 El Valle),
Lesser Nighthawk (1 Chagres), Common Paraque (heard Hotel Radisson Summit, 1 Fortuna),
Dusky Nightjar (2 Cerro Santiago, perhaps 3 or 4 in the vicinity),
[Common (Northern) Potoo – Metiti],
Black Swift (a few Burbayar), Chesnut-collared Swift (small numbers western highlands),
White-collared Swift (common western highlands),
Vaux's Swift ( a few western highlands), Short-tailed Swift (several Chagres, Burbayar),
Band-rumped Swift (common eastern lowlands), Costa Rican Swift (a couple Boquete),
Grey-rumped Swift (a few Fortuna), Lesser Swallow-tailed Swift (widespread, sparse),
White-necked Jacobin (several Chagres, Torti), Rufous-breasted Hermit (a couple Torti),
Green Hermit (common Cerro Pirre and western highlands),
Long-billed Hermit (small numbers near Yaviza, Rancho Frio),
White-whiskered Hermit (a couple Cerro Pirre ridge), Pale-bellied Hermit (2 Rancho Frio),
Stripe-throated Hermit (widespread in lowlands, fairly common),
Green-fronted Lancebill (1 Cerro Pirre, 1 Fortuna),
Brown Violetear (1 or 2 Cerro Pirre ridge, a few western highlands), Green Violetear (2 Boquete),
Tooth-billed Hummingbird (2 or 3 Cerro Pirre upper slopes),
Purple-crowned Fairy (small numbers Cerro Pirre), Black-throated Mango (a couple Torti),
Veraguan Mango (1 Las Lajas),
Green-crowned Brilliant (fairly common, widespread),
Magnificent Hummingbird (common Volcan Buru), Fiery-throated Hummingbird (3 Volcan Buru),
Long-billed Starthroat (a few Torti, Burbayar),
White-bellied Mountain-Gem (common Fortuna),
Purple-throated Mountain-Gem (common Fortuna, Cerro Santiago),
White-throated Mountain-Gem (common Volcan Buru),
Magenta-throated Woodstar (1 Volcan Buru), Volcano Hummingbird (common Volcan Buru),
Scintillant Hummingbird (fairly common Volcan area; nesting in hotel grounds),
Garden Emerald (a few El Valle, Volcan, Las Lajas),
Violet-headed Hummingbird (1 Cerro Pirre upper slopes, 1 Fortuna),
Scaly-breasted Hummingbird (small numbers Torti, near Yaviza, Burbayar),
Violet Sabrewing (1 or 2 Boquete), Stripe-tailed Hummingbird (1 Boquete),
Black-bellied Hummingbird (small numbers Fortuna),
White-tailed Emerald (several western highlands),
Snowcap ( 3 or 4 Fortuna),
White-vented Plumeleteer (1 El Valle), Bronze-tailed Plumeleteer (1 El Valle, 1 Burbayar),
Crowned Woodnymph (fairy common, widespread),
Blue-chested Hummingbird (common Burbayar, a few Ranch Frio),
Charming Hummingbird (a couple Volcan),
Snowy-bellied Hummingbird (small numbers, widespread in lowlands),
Rufous-tailed Hummingbird (widespread),
Pirre Hummbird (small numbers Cerro Pirre ridge & higher slopes),
Sapphire-throated Hummingbird (uncommon, widespread in lowlands),
Violet-bellied Hummingbird (1 Chagres),
Lattice-tailed Trogon (1 Fortuna),
Slaty-tailed Trogon (several western lowlands), White-tailed Trogon (a few Pipeline Rd, Torti),
Gartered Trogon (common Chagres, Pipeline Rd),
Black-throated Trogon (uncommon western lowlands),
Collared Trogon (several Cerro Pirre and westerm highlands),
Orange-bellied Trogon (1 Boquete), Resplendent Quetzel (several Volcan Buru),
Tody Motmot (2 or 3 Cerro Pirre foothills),
Whooping Motmot (several western lowlands), Blue-crowned Motmot (1 El Valle),
Rufous Motmot (1 Burbayar), Broad-billed Motmot (a few Burbayar, Rancho Frio, Pipeline Rd),
Ringed Kingfisher, Belted Kingfisher, Amazon Kingfisher, Green Kingfisher,
Barred Puffbird (2 near Yaviza),
[White-necked Puffbird – Rancho Frio, El Valle],
Black-breasted Puffbird (1 Pipeline Rd), Pied Puffbird (2 Rancho Frio, heard Torti),
White-whiskered Puffbird (several Chagres, Pipeline Rd, Rancho Frio),
Grey-cheeked Nunlet (2 near Yaviza),
White-fronted Nunbird (1 near Yaviza),
[Rufous-tailed Jacamar – near Yaviza], Great Jacamar (a couple Rancho Frio),
Spot-crowned Barbet (small numbers near Yaviza, Rancho Frio, Cerro Pirre mid-elevation),
Red-headed Barbet (a few Burbayar, El Valle), Prong-billed Barbet (common western highlands),
Emerald Toucanet (Blue-throated caeruleogularis - fairly common western highlands; Violet-throated cognatus – 1 Cerro Pirre upper slopes),
Collared Aracari (common eastern lowlands), Fiery-billed Aracari (a few Volcan).
Yellow-eared Toucanet (small numbers Cerro Pirre foothills & mid-elevation),
Keel-billed Toucan (common, widespread). Chesnut-mandibled Toucan (widespread, uncommon).
Olivaceous Piculet (1 Rancho Frio, 1 Volcan), Acorn Woodpecker (common Volcan Buru),
Black-cheeked Woodpecker (2 Pipeline Rd), Red-crowned Woodpecker (widespread),
Hairy Woodpecker (small numbers western highlands), Smoky-brown Woodpecker (1 Volcan),
Red-rumped Woodpecker (a couple Rancho Frio),
Stripe-cheeked Woodpecker (2 Cerro Pirre mid-elevation),
Golden-olive Woodpecker (a few western highlands), Spot-breasted Woodpecker (3-4 Rancho Frio),
Cinnamon Woodpecker (uncommon eastern lowlands),_Lineated Woodpecker (widespread),
Crimson-bellied Woodpecker (small numbers Rancho Frio, Burbayar),
Crimson-crested Woodpecker (widespread, uncommon),
Barred Forest-Falcon (1 El Valle, heard Fortuna), Slaty-backed Forest-Falcon (1 Pipeline Rd),
Red-throated Caracara (a few Rancho Frio), Crested Caracara, Yellow-headed Caracara,
Laughing Falcon (widespread, uncommon), American Kestrel, Peregrine Falcon (1 near Las Lajas),
Aplomado Falcon (1 en route to Cerro Santiago), Bat Falcon (2 Radisson Summit Hotel, 2 Burbayar),
Sulphur-winged Parakeet (2 flocks Boquete),
Crimson-fronted Parakeet (uncommon western highlands), Brown-throated Parakeet (a few Boquete),
Great Green Macaw (a few Rancho Frio, Cerro Pirre, Burbayar),
Red-and-green Macaw (several Cerro Pirre mid-elevation),
Blue-and-yellow Macaw (a few Ranch Frio),
[Barred Parakeet – Volcan Buru], Orange-chinned Parakeet (widespread),_
Spectacled Parakeet (uncommon Rancho Frio, from boat to El Real),
Blue-fronted Parrotlet (feeding flock Cerro Pirre upper slopes),
Brown-hooded Parrot (small numbers eastern lowlands), Blue-headed Parrot (widespread),
Red-lored Parrot (widespread), Mealy Parrot (fairly common),
Sapayoa (a trip highlight: total 12-14 at Rancho Frio, Cerro Pirre mid-elevation, Burbayar),
Fasciated Antshrike (uncommon eastern lowlands), Great Antshrike (1 Rancho Frio, 1 Boquete),
Barred Antshrike (2 Torti), [Black-hooded Antshrike – Las Lajas],
Black Antshrike (2 Rio Mono, heard Yaviza),
Western Slaty-Antshrike (common eastern lowlands), {Speckled Antshrike (2 Burbayar)},
Russet Antshrike (small numbers Cerro Pirre, Burbayar),
Plain Antvireo (1 Cerro Pirre, 2 Boquete), Spot-crowned Antvireo (common eastern foothills),
Moustached Antwren (a few Pipeline Rd, Torti),
Pacific Antwren (2 Torti),
White-flanked Antwren (widespread in lowlands, sparse),
Checker-throated Antwren (widespread, uncommon),
Slaty Antwren (a few Burbayar, El Valle), Rufous-winged Antwren (1 Pipeline Rd, 1 Rancho Frio),
Dot-winged Antwren (uncommon eastern lowlands), Dusky Antbird (fairly common eastern lowlands),
White-bellied Antbird (1 year Yaviza others heard, heard Rancho Frio),
Chesnut-backed Antbird (common eastern lowlands),
Dull-mantled Antbird (small numbers Cerro Pirre foothills & mid-elevation, Burbayar),
Immaculate Antbird (a few Cerro Pirre mid-elevation),
Spotted Antbird (small numbers Pipeline Rd, Rancho Frio, Burbayar),
Wing-banded Antbird (2 Cerro Pirre mid-elevation was a trip highlight),
Bicoloured Antbird (fairly common eastern lowlands, mainly heard),
Ocellated Antbird (several Burbayar),
Black-crowned Antpitta (a trip highlight: a few Cerro Pirre mid-elevation, Rancho Frio, Burbayar),
Streak-breasted Antpitta (1 Pipeline Rd, 1 Rancho Frio),
Ochre-breasted Antpitta (1 Fortuna),
{Choco Tapaculo – 2-3 Cerro Pirre ridge},
Silvery-fronted Tapaculo (several Volcan Buru, heard Cerro Santiago),
Black-faced Ant-thrush (1 El Valle; heard Pipeline Rd, Chagres),
Black-headed Ant-thrush (1 Fortuna),
Scaly-throated Leaftosser (1 Chagre, 1 Cerro Pirre mid-elevation),
Plain-brown Woodcreeper (fairly common eastern lowlands),
Olivaceous Woodcreeper (1 Chagre), Wedge-billed Woodcreeper (fairly common, widespread),
Northern Barred-Woodcreeper (1 Burbayar), Cocoa Woodcreeper (common eastern lowlands),
Spotted Woodcreeper (a few Burbayar, El Valle), Straight-billed Woodcreeper (2 Burbayar),
Brown-billed Scythebill (2 Cerro Pirre mid-elevation, 1 Fortuna),
Streak-headed Woodcreeper (sparse Cerro Pirre, western highlands),
Plain Xenops (widespread), [Streaked Xenops – Torti],
[Sharp-tailed Streamcreeper – Cerro Pirre mid-elevation],
Buffy Tuftedcheek (2 Volcan Buru, 1 Cerro Santiago), [Scaly-throated Foliage-gleaner – Boquete],
Slaty-winged Foliage-gleaner (small numbers Rancho Frio),
Lineated Foliage-Gleaner (a few Cerro Pirre upper slopes, 1 Volcan Buru),
Striped (Western) Woodhaunter (1 Rancho Frio, 1 Burbayar),
Buff-throated Foliage-gleaner (2 Rancho Frio), Ruddy Foliage-gleaner (1 Boquete),
Streak-breasted Treehunter (1 Boquete, several Cerro Santiago),
Spotted Barbtail (a few western highlands), {Beautiful Treerunner (1 Cerro Pirre ridge)},
Ruddy Treerunner (small numbers western highlands),
Double-banded Greytail (pairs Torti, Rancho Frio),
Red-faced Spinetail (fairly common western highlands),
Pale-breasted Spinetail (a couple Volcan), Slaty Spinetail (1 Cerro Pirre, 1 Boquete),
Brown-capped Tyrannulet (sparse eastern lowlands),
Southern Beardless-Tyrannulet (1 Las Lajas), Yellow Tyrannulet (a few El Valle, Volcan),
Yellow-crowned Tyrannulet (singles Rancho Frio, Burbayar),
Forest Elaenia (uncommon eastern lowlands), Grey Elaenia (uncommon eastern lowlands),
Greenish Elaenia (a few Chagres, Boquete), Yellow-bellied Elaenia (widespread),
Lesser Elaenia (singles Chagres, Volcan), Mountain Elaenia (several western highlands),
Torrent Tyrannulet (1 Boquete), Olive-striped Flycatcher (small numbers western highlands),
Ochre-bellied Flycatcher (sparse, widespread), Slaty-capped Flycatcher (1 Fortuna),
Yellow-green Tyrannulet (2 Chagres, 2 Cerro Pirre mid-elevation),
Rufous-browed Tyrannulet (1 Fortuna),
Rough-legged (White-fronted) Tyrannulet (a few Volcan Buru, Boquete),
Paltry Tyrannulet (small numbers Cerro Pirre, western highlands),
Black-capped Pygmy-Tyrant (1 Pipeline Rd),
Scale-crested Pygmy-Tyrant (widespread in foothills, sparse),
Southern Bentbill (singles Pipeline Rd, Rancho Frio),
Slate-headed Tody-Flycatcher (1 Chagres), Common Tody-Flycatcher (sparse),
Black-headed Tody-Flycatcher (2 Chagres), Brownish Twistwing (1 Pipeline Rd),
Eye-ringed Flatbill (a few Cerro Pirre mid-elevation, Volcan, Fortuna).
Olivaceous Flatbill (sparse eastern lowlands, El Valle),
Yellow-olive Flycatcher (a few Pipeline Rd, El Valle, Volcan),
Yellow-margined Flycatcher (uncommon, widespread), Yellow-breasted Flycatcher (2 Rancho Frio),
White-throated Spadebill (a few El Valle, western highlands),
Golden-crowned Spadebill (1 Rancho Frio, another heard),
Ruddy-tailed Flycatcher (fairly common lowlands, foothills),
Sulphur-rumped Flycatcher (small numbers Rancho Frio, Burbayar),
Common Tufted-Flycatcher (a few western highlands), Olive-sided Flycatcher (widespread, sparse),
Dark Pewee (a few Boquete),
Ochraceous Pewee (1 Volcan Buru),
Western Wood-Pewee (singles Rancho Frio, El Valle, Cerro Santiago),
Eastern Wood-Pewee (common, widespread), Tropical Peewee (2 Hotel Radisson Summit),
Yellow-bellied Flycatcher (a few Volcan, Boquete), Acadian Flycatcher (widespread, uncommon),
{Alder Flycatcher (1 El Valle)}, Willow Flycatcher (sparse), Least Flycatcher (1 Las Lajas),
Yellowish Flycatcher (singles Volcan Buru, Cerro Santiago),
Black-capped Flycatcher (2 Volcan Buru), Black Phoebe (sparse western highlands),
Pied Water-Tyrant (several east of Metiti), Long-tailed Tyrant (2 east of Metiti),
Bright-rumped Attila (sparse but widespread eastern lowlands),
Choco Sirystes (2 Rancho Frio),
Rufous Mourner (1 Pipeline Rd), Dusky-capped Flycatcher (widespread in small numbers),
Panama Flycatcher (singles Chagre, Burbayar), Great Crested Flycatcher (a few Burbayar),
Lesser Kiskadee (2 east of Metiti), Great Kiskadee, Boat-billed Flycatcher,
Rusty-margined Flycatcher (sparse eastern lowlands), Social Flycatcher,
Grey-capped Flycatcher (singles Burbayar, Boquete),
Golden-bellied Flycatcher (singles Volcan Buru, Cerro Santiago),
Streaked Flycatcher (widespread), Sulphur-bellied Flycatcher (singles Chagres, Torti),
Piratic Flycatcher (a few Chagres, Pan-American Rd), Tropical Kingbird,
Eastern Kingbird (small numbers eastern lowlands), Fork-tailed Flycatcher (widespread, sparse),
Russet-winged Schiffornis (1 Pipeline Rd),
{Northern Schiffornis (1 El Valle, others heard)},
Speckled Mourner (1 Rancho Frio),
Barred Becard (a few western highlands), Cinnamon Becard (a few Pan-American Rd),
White-winged Becard (sparse eastern lowlands), Masked Tityra (widespread),
Black-crowned Tityra (a few Rancho Frio, Burbayar),
Purple-throated Fruitcrow (fairly common Pipleine Rd),
Blue Cotinga (1 Chagres, several Rancho Frio),
Rufous Piha (a few Rancho Frio, Burbayar), Green Manakin (1 Rancho Frio, 2 Burbayar),
Orange-collared Manakin (2 Volcan),
Golden-collared Manakin (fairly common eastern lowlands),
White-ruffed Manakin (small numbers Cerro Pirre mid-elevation, Burbayar, El Valle, Boquete),
Lance-tailed Manakin (a few Chagres, Boquete), Blue-crowned Manakin (widespread, uncommon),
White-crowned Manakin (1 Fortuna), Golden-headed Manakin (several Rancho Frio),
Red-capped Manakin (a few Hotel Radisson Summit, Pipeline Rd, Burbayar),
Sharpbill (2 Cerro Pirre mid-elevation),
Yellow-throated Vireo (2 Boquete), Yellow-winged Vireo (several Volcan Buru, Boquete),
Brown-capped Vireo (fairly common western highlands),
Philadelphia Vireo (1 Boquete),
Red-eyed Vireo, Yellow-green Vireo (a few Burbayar, Boquete),
Scrub Greenlet (1 Hotel Radisson Summit), Tawny-crowned Greenlet (2 Chagres),
Golden-fronted Greenlet (1 Chagres), Lesser Greenlet (widespread),
Green Shrike-Vireo (1 Pipeline Rd, heard Chagres),
[Yellow-browed Shrike-Vireo - Cerro Pirre mid-elevation],
Rufous-browed Peppershrike (a few Volcan Buru, Boquete),
Silvery-throated Jay (flock Volcan Buru),
Black-chested Jay (widespread, sparse),
Grey-breasted Martin, Brown-chested Martin (a few Burbayar),
Tree Swallow (2 PipelineRd), Mangrove Swallow (widespread in small numbers),
Blue-and-white Swallow (common western highlands),
Southern Rough-winged Swallow (common eastern lowlands),
Bank Swallow, Cliff Swallow, Barn Swallow, Cave Swallow (2 Fortuna),
Southern Nightingale-Wren (widespread, mostly heard),
House Wren, Ochraceous Wren (several western highlands),
White-headed Wren (a few Rancho Frio), Rufous-breasted Wren (sparse in lowlands),
Sooty-headed Wren (several Rancho Frio ridge),
Black-bellied Wren (fairly common eastern lowlands),
Rufous-and-white Wren (2 Hotel Radisson Summit),
Stripe-throated Wren (several Rancho Frio, heard Burbayar),
[Stripe-breasted Wren – Fortuna], Plain Wren (several El Valle, Boquete),
Bay Wren (widespread), Riverside Wren (a few Volcan),
Buff-breasted Wren (uncommon Pan-American Rd),
White-breasted Wood-Wren (common, widespread),
Grey-breasted Wood-Wren (common in highlands),
Song Wren (singles Pipeline Rd, El Valle; others heard),
American Dipper (1 Boquete),
Tawny-faced Gnatwren (widespread in lowlands, sparse), Long-billed Gnatwren (2 Chagres),
Tropical Gnatcatcher (widespread), Slate-throated Gnatcatcher (a few Rancho Frio),
Black-capped Donacobius (a few from boat to El Real),
Black-faced Solitaire (common western highlands),
Varied Solitaire (several Cerro Pirre ridge, upper slopes),
Black-billed Nightingale-Thrush (2 Volcan Buru),
Orange-billed Nightingale-Thrush (common western highlands),
Slaty-backed Nightingale-Thrush (small numbers western highlands),
Ruddy-capped Nightingale-Thrush (a few Volcan Buru, Boquete),
[Black-headed Nightingale-Thrush – Fortuna],
Veery (a few Chagre, Pipeline Rd), Grey-cheeked Thrush (1 Cerro Pirre),
Swainson's Thrush (widespread), Mountain Thrush (common western highlands),
Pale-vented Thrush (a few El Valle, Fortuna, Cerro Santiago),
Clay-coloured Thrush (widespread), White-throated Thrush (1 Boquete), Tropical Mockingbird,
Black-and-yellow Silky-Flycatcher (several Volcan Buru, Cerro Santiago),
Long-tailed Silky-Flycatcher (a few Volcan Buru),
Ovenbird (2 Fortuna), Northern Waterthrush (common),
Golden-winged Warbler (uncommon El Valle, western highlands),
Black-and-white Warbler (fairly common El Valle, western highlands),
Flame-throated Warbler (sparse western highlands), Tennessee Warbler (widespread),
Connecticut Warbler (1 Hotel Radisson Summit, 1 Chagre),
Masked (Chiriqui) Yellowthroat (2 Volcan), Olive-crowned Yellowthroat (1 Fortuna),
Mourning Warbler (widespread, uncommon), American Redstart (1 Boquete),
Tropical Parula (a few western highlands), Bay-breasted Warbler (2 Fortuna),
Blackburnian Warbler (widespread), Yellow Warbler (widespread),
Chesnut-sided Warbler (several western highlands),
Townsend's Warbler (1 Volcan Buru), Golden-cheeked Warbler (1 Volcan Buru),
Black-throated Green Warbler (several western highlands),
Buff-rumped Warbler (a few Rancho Frio, Burbayar),
Rufous-capped Warbler (small numbers El Valle, western highlands),
Black-cheeked Warbler (common Volcan Buru, Cerro Santiago),
{Pirre Warbler (a few Cerro Pirre ridge)}, Golden-crowned Warbler (several Boquete, Cerro Santiago),
Three-striped Warbler (a few Boquete, Fortuna), Canada Warbler (widespread),
Wilson's Warbler (common western highlands),
Slate-throated Redstart (common Cerro Pirre, western highlands),
Collared Redstart (common western highlands),
Zeledonia (Wrenthrush) (3 Volcan Buru),
Bananaquit, Yellow-backed Tanager (2 Rancho Frio),
Black-and-yellow Tanager (fairly common Cerro Pirre mid-elevation, Fortuna),
Rosy Thrush-Tanager (1 Chagre, heard Las Lagas),
Dusky-faced Tanager (several Fortuna), Grey-headed Tanager (2 Pipeline Rd),
Sulphur-rumped Tanager (several Burbayar),
Scarlet-browed Tanager (a few Cerro Pirre mid-elevation),
White-shouldered Tanager (common in lowlands),
Tawny-crested Tanager (a few Burbayar, El Valle),
White-lined Tanager (several Chagre, Pipeline Rd),
Crimson-backed Tanager (common eastern lowlands, El Valle),
Cherrie's Tanager (small numbers Volcan, Boquete),
Flame-rumped Tanager (a few Burbayar, El Valle),
Blue-grey Tanager, Palm Tanager,
Blue-and-gold Tanager (fairly common Fortuna),
Grey-and-gold Tanager (a few Cerro Pirre), Golden-hooded Tanager (widespread, sparse),
Speckled Tanager (widespread, uncommon), {Green-naped Tanager (2 Cerro Pirre ridge)},
Spangle-cheeked Tanager (2 Cerro Santiago),
Plain-coloured Tanager (uncommon eastern lowlands, El Valle),
Rufous-winged Tanager (a few Burbayar), Bay-headed Tanager (widespread, sparse),
Emerald Tanager (uncommon, widespread),
Silver-throated Tanager (fairly common, widespread),
Scarlet-thighed Dacnis (sparse western highlands), Blue Dacnis (fairly common, widespread),
Viridian Dacnis (1 Rancho Frio),
Green Honeycreeper (uncommon, widespread), Shining Honeycreeper (a few Burbayar, Volcan),
Red-legged Honeycreeper (several Rancho Frio, Volcan),
Streaked Saltator (a few Torti, El Valle), Buff-throated Saltator (widespread),
Slate-coloured Grosbeak (several Chagre, Cerro Pirre, Burbayar),
Blue-black Grassquit, Yellow-faced Grassquit, Variable Seedeater,
Yellow-bellied Seedeater (uncommon Chagre, western highlands),
Ruddy-breasted Seedeater (a few east of Metiti),
Lesser (Thick-billed) Seed-Finch (singles Chagre, Torti),
Slaty Flowerpiercer (a few western highlands), Saffron Finch (common Hotel Radisson Summit),
Yellow-thighed Finch (a few Volcan Buru, Boquete),
Yellow-green Finch (common Cerro Santiago),
Large-footed Finch (several Volcan Buru), Orange-billed Sparrow (1 Chagre),
Chesnut-capped Brush-Finch (sparse El Valle, western highlands),
Costa Rican Brush-Finch (a few Volcan),
Black-striped Sparrow (a few El Valle), Rufous-collared Sparrow (uncommon western highlands),
White-naped Brush-Finch (sparse Volcan Buru, Cerro Santiago),
Common Bush-Tanager (fairly common El Valle punctulatus, western highlands novicius),
Pirre Bush-Tanager (fairly common Cerro Pirre ridge, higher slopes),
Sooty-capped Bush-Tanager (several Volcan Buru, Cerro Santiago).
Yellow-throated Bush-Tanager (2 Fortuna), Hepatic Tanager (widespread, sparse),
Summer Tanager (common), Scarlet Tanager (a few Rancho Frio, El Valle),
Flame-coloured Tanager (uncommon western highlands),
White-winged Tanager (a few Volcan, Boquete),
Red-crowned Ant-Tanager (a few El Valle, Volcan),
Red-throated Ant-Tanager (uncommon Chagre, Pipeline Rd, El Valle),
Olive (Carmiol's) Tanager (a few Burbayar),
Lemon-spectacled Tanager (fairly common Rancho Frio, Cerro Pirre mid-elevation),
Black-faced Grosbeak (2 Fortuna), [Yellow-green Grosbeak - Cerro Pirre],
Black-thighed Grosbeak (2 Volcan Buru), Rose-breasted Grosbeak (a few Volcan Buru),
Blue-black Grosbeak (sparse eastern lowlands), Dickcissel (a few Pipeline Rd, Pan-American Rd),
Bobolink (a few Pan-American Rd), Yellow-hooded Blackbird (several east of Metiti),
Red-breasted Blackbird ( a few east of Metiti), Eastern Meadowlark (sparse western highlands),
Melodious Blackbird (a few Volcan Buru), Great-tailed Grackle,
Giant Cowbird (widespread, sparse), Orchard Oriole (a few Pan American Rd),
Yellow-backed Oriole (a few Chagre, Rancho Frio),
[Orange-crowned Oriole – Pan American Rd], [Yellow-billed Cacique – Boquete],
Scarlet-rumped Cacique (a few Pipeline Rd, Rancho Frio, Fortuna),
Yellow-rumped Cacique (a few Hotel Radisson Summit, Rancho Frio),
Crested Oropendola (sparse Burbayar, western lowlands),
Chesnut-headed Oropendola (fairly common eastern lowlands, El Valle),
Black Oropendola (2 year Yaviza; several from boat to El Real),
Yellow-crowned Euphonia (sparse, widespread), Elegant Euphonia (several western highlands),
Thick-billed Euphonia (fairly common, widespread),
Fulvous-vented Euphonia (several Pipeline Rd, Rancho Frio, Burbayar),
Spot-crowned Euphonia (a few El Valle), Olive-backed Euphonia (2 Boquete),
White-vented Euphonia (1 Rancho Frio), Tawny-capped Euphonia (widespread, sparse),
Orange-bellied Euphonia (fairly common Cerro Pirre),
Golden-browed Chlorophonia (several western highlands),
Yellow-bellied Siskin (a few Volcan), Lesser Goldfinch (widespread, sparse), House Sparrow.


Mantled Howler (Hotel Radisson Summit, Pipeline Rd), White-faced Capuchin,
Geoffroy's Tamarin (several near Yaviza, Burbayar),
Brown Three-toed Sloth (seen several times),
{Hoffmann's Two-toed Sloth – 1 Hotel Radisson Summit},
Central American Agouti, Nine-banded Armadillo,
Variegated Squirrel, Red-tailed Squirrel,
Western Pygmy-Squirrel Microsciurus mimulus, Alfaro's Pygmy Squirrel M. alfari,
Capybara (several Hotel Radisson Summit), White-tailed Deer.